Department of Plant Pathology and Microbiology, University of California, Riverside, CA, 92521, USA. Center for Plant Cell Biology, University of California, Riverside, CA, 92521, USA. Department of Plant Pathology and Microbiology, University of California, Riverside, CA, 92521, USA. wenbo.ma@ucr.edu. Center for Plant Cell Biology, University of California, Riverside, CA, 92521, USA. wenbo.ma@ucr.edu.